Cultivating Carnivorous Plants (2015)
DROSERA
Drosera regia with prey.12
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The genus Drosera must be the most
diverse of all the genera that comprise the
carnivorous plants. Sticky-leaved sundews
grow from the edges of the Arctic Circle
to the Equator and south to the Tierra del
Fuego. Some are giants, growing as large
as small shrubs; others are pygmies, only
growing as large as a small coin. Some live
in tropical rainforests while others live in
deserts. There are few places where
sundews cannot be found, and that makes
them perfect for beginners.
Although
incredibly
diverse,
sundews are characterized by leaves
covered in stalked mucilaginous glands,
often called tentacles, that produce a
sticky substance, which binds insects and
other small prey to the leaves. As the prey
struggles, more sticky mucus is produced,
binding the prey tighter. As this occurs,
the plant secretes digestive enzymes
(esterase, peroxidase, phosphatase, and
protease) that dissolve the prey’s soft,
fleshy parts. The resulting “soup” is then
Drosera capensis x spathulata, an easy and popular hybrid.
absorbed by the leaf and used as fertilizer
for the plant.
Unlike Byblis, Drosophyllum, or Roridula, all of which are phonologically similar to Drosera, all species of sundew have tentacles that move in response to contact with prey. This response allows the plant to gradually
move the insect towards the center of the leaf, allowing more tentacles to come in contact with the prey. This
improved contact allows the plant to more effectively digest its prey. Additionally, many species--although
not all--also move their leaves in response to contact with prey. Typically, this results in the leaf slowly
wrapping itself around the prey.
Although diverse, most sundews can be divided into the following horticultural categories:
✵
Temperate: These species require a cold winter dormancy period. In nature, they will often
experience a period of freezing temperatures. In order to survive those cold temperatures, they will
form a tight cluster of unfurled leaves, called a hibernaculum. All of these species are native to areas
that can experience snowfall.
✵
Tropical & Subtropical: These species will continue to grow year-round in cultivation without
needing a dormancy period. In nature, the tropical varieties grow year-round in nearly uniform
climatic conditions, but the subtropical species experience a cool winter.
✵
Highland Tropical: These species will grow year-round in cultivation without needing a dormancy
period. In nature, they grow in highland areas of the tropics, or other areas that experience nearly
uniform climatic conditions, with temperature falls at night and daytime temperatures that remain
cool. These are the species that can be found, e.g. on the Tepuis of Venezuela.
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CULTIVATING CARNIVOROUS PLANTS
✵
Pygmy: These miniature species of sundew are mostly native to Australia (one species, Drosera
meristocaulis, is native to South America), and naturally reproduce in two different ways - through
seed and production of gemmae. Gemmae are small "buds" consisting of a plantlet and food
reservoir that appear during the wet season. As rain hits the gemmae, which form around the crown
of the plant, they are hurled from the plant and form new plants.
✵
Tuberous: These Australian species form underground tubers to survive the dry summers of their
habitat.
✵
Petiolaris-Complex: These Australian species are similar to Drosophyllum in that they are well
adapted for living in a dry native habitat. Most have "hairy" petioles that allow the plant to capture
morning fog.
In Cultivation
Given the diverse nature of the horticultural categories of Drosera, the best way to discuss their
cultivation is by discussing each of the horticultural categories individually. This follows:
Temperate Drosera
Drosera rotundifolia in the wild.
The most famous temperate Drosera and, arguably, the most famous Drosera, is Drosera rotundifolia. Native
to the northern hemisphere, this species is one of the most widespread of all the Drosera, and the carnivore
most studied by Charles Darwin and a host of scientists after him. Surprisingly, despite its fame and
widespread distribution, Drosera rotundifolia, like many temperate Drosera, is not as common in cultivation as
some tropical and subtropical species.
With regard to cultivation requirements, temperate Drosera are somewhat more difficult to cultivate than
subtropical Drosera because they require a cold dormancy period of several months. This is not an issue for
growers in colder climates, where growers can leave the plants outside year round and expect that they will
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return in the spring. For growers in warmer
climates, plans must be made regarding how to
chill the plants in winter for several months in
a manner that will not result in fungus
overwhelming the plants and killing them.
Thanks to the fact that many temperate Drosera
form “winter buds” called hibernacula,
oftentimes, growers will have success in
overwintering the plants by placing them in a
refrigerator after dusting them with fungicide.
D. binata - Complex
The unmistakable Forked Sundew,
Drosera binata, is native to Australia, New
Zealand, and Tasmania. As with the Drosera
filiformis Complex, many members of the
Drosera binata Complex are temperate, while
others are subtropical. The major difference
between temperate and subtropical individuals
is that temperate individuals will go dormant in
cool weather, dying down to their roots,
whereas subtropical specimens will continue
growing year round, unless killed back by frost.
As with other “complexes,” Drosera binata
has a number of distinctive forms. The most
common are the ‘T-Form’, which consists of
Drosera binata ‘Extrema’
short (6 inch (15 cm)) single-forked leaves,
‘Extrema’, which consists of medium-height (12 inch (30 cm)) leaves forked up to 40 times, ‘Dichotoma
Giant', which has leaves up to 30 inches (76 cm) forked, usually, less than a half-dozen times, and ‘Marston
Dragon’, a cultivar with leaves up to 12 inches (30 cm) forked twice.
D. filiformis - Complex
The most famous of the “thread-leaf” sundews, Drosera filiformis is
native to several, distinct locations along the east coast of North America.
These locations include, Nova Scotia, Cape Cod, Long Island/New Jersey,
Cape Fear area of North Carolina, and the Panhandle of Florida. The
distribution of Drosera filiformis to these distinct locations is likely due to
migrating birds along the east coast gathering seed from populations in
seed on their muddied feet then flying to a new location. This theory is
supported by the fact that the species has been spontaneously reported in
other locations ( e.g. Rhode Island, Connecticut, and Maryland) where the
species was not previously known. (The two New England locations are
now extinct). It is unclear why the spread of the species to new locations
has not resulted in a continuous distribution throughout its range. Plants Drosera filiformis ssp. filiformis
from these locations are known as Drosera filiformis ssp. filiformis.
in Massachusetts.
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CULTIVATING CARNIVOROUS PLANTS
In Florida, another subspecies
of Drosera filiformis exists. This
subspecies, known as Drosera filiformis
ssp. trayci, has thick green tentacles
and lacks red coloration. It is also
much taller than Drosera filiformis ssp.
filiformis, and is often found growing
alongside Sarracenia leucophylla or in
pure white sands. Although it has
been argued that Drosera filiformis ssp.
trayci is a separate species, there has
not been enough evidence presented
to distinguish Drosera filiformis ssp.
trayci as a separate species. Drosera
filiformis ssp. tracyi differs from Drosera
filiformis ssp. filiformis in the following
Drosera filiformis ssp. trayci
respects:
Character
D. filiformis ssp. filiformis
D. filiformis ssp. tracyi
Leaf Length
3 - 10 inches (8 - 26 cm)
12 - 20 inches (30 - 51 cm)
Scape Length
2.4 - 10 inches (6 - 26 cm)
10 - 24 in (26 - 61 cm)
Color of Glandular Hair
Red
Clear
Color of Plant
Reddish
Pale Green
For Drosera filiformis ssp. filiformis from the northern
locations, i.e. those not from Florida, cultivation follows the
traditional
temperate
Drosera
pattern.
In
winter,
hibernaculum form and the plant must be subjected to cold
winter temperatures. For plants from Florida, often known
as ‘Florida Giant’, ‘Florida Red’, ‘Giant Red’, or ‘Red’,
hibernaculum are, occasionally, formed for a month-or-so
upon near-freezing temperatures being reached, but the
plants begin growing immediately upon temperatures
warming, a death sentence for more northerly climates,
where freezing temperatures will kill the new growth.
A comprehensive genetic study of the different
populations of Drosera filiformis ssp. filiformis has never been
undertaken. Such a study would probably reveal genetic
differences significant enough to merit a new classification
of this species. Already, there is significant evidence that
the Florida variant is different from the other populations.
Currently, most Drosera filiformis ssp. filiformis in
cultivation are from New Jersey, except for the ‘Florida
Drosera filiformis ssp. filiformis ‘Florida Red’
Red’ plants. The plants from New Jersey are larger and less has pink flowers, much like the other variants of
cold tolerant than the ones from more northern locales.
Drosera filiformis ssp. filiformis.
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D. rotundifolia
Drosera rotundifolia is the
quintessential temperate Drosera
and a must have for any grower
interested in temperate Drosera.
It is a remarkably easy species
to grow, if given a cold winter
dormancy. In good conditions,
one plant can produce upwards
of a thousand seeds per season!
Despite growing around
the northern hemisphere, the
first anthocyanin-free specimen
( f.
viridescens)
of
Drosera
rotundifolia was found in 2006
near Iga City, Mie Prefecture,
Kinki District, Japan. It is
identical in all aspects to
regular
Drosera
rotundifolia,
except that it does not possess
Drosera rotundifolia growing in a bog in New Hampshire.
any red pigment.
My standard growing guide for temperate Drosera follows:
✵
Media: Temperate Drosera are often grown in a mix of one part sphagnum peat to one part sand or
perlite, but some, such as Drosera linearis appear to require a less acidic media. A mix of one part
sphagnum peat to alkaline clay works well for the species that prefer a less acidic media.
✵
Moisture: Watering via the tray method, and always keeping some water in the tray is probably the
easiest way to meet the moisture requirements of temperate Drosera during the growing season.
During dormancy, soil should remain moist but not soaking in order to prevent rot.
✵
Humidity: Temperate Drosera are not particularly picky about humidity, but, in nature, typically
experience somewhat higher ambient humidity during the growing season than in dormancy.
✵
Pot Size: Pots should be larger than the plant in diameter, but most Drosera are not particular about
pot size.
✵
Feeding: Feeding temperate Drosera will encourage growth. The more insects caught or orchid
fertilizer spritzed on them, the better they will grow. Typically, I use half-strength orchid fertilizer
applied to the leaves of plants grown indoors. Plants grown outdoors will capture enough small
insects on their own that fertilizer is not strictly necessary.
✵
Temperature: Temperate species can tolerate a wide range of above-freezing temperatures during the
growing season, but need to experience a cold, but not necessarily freezing, dormancy period lasting
several months.
✵
Dormancy: Dormancy is required for temperate Drosera. In cold climates, plants can be left outdoors,
but in warmer climates, plants should be dug up after forming hibernaculum in mid-autumn. These
hibernaculum should be wrapped in a damp paper towel, placed in a plastic bag, and put in a
refrigerator for several months.
✵
Propagation: Temperate Drosera are most easily propagated through seed, but can also be propagated
through leaf cuttings.
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CULTIVATING CARNIVOROUS PLANTS
o
Seed: The easiest way to propagate temperate Drosera is to collect their copious seed. Seed is
mature when the pods dry, turn black, and begin to split. The seed requires a cool
stratification period of at least four weeks. The best way to do this is to sow the seed on a
damp, suitable medium, place plastic wrap over the medium, and place in the refrigerator for
a month. The seed should then be removed and placed in a well-lit area. Seedlings should
appear within weeks.
o
Leaf Cuttings: Leaf cuttings are a remarkably easy method of propagation. As with other
Drosera, cut the leaf from its base and place on an appropriate medium. In a few weeks, a
small plantlet should appear near one of the tentacles. It is beneficial to give the cutting
higher humidity than the parent plants.
Tropical & Subtropical Drosera
The tropical and subtropical Drosera group includes a wide
variety of easy to grow species including the often-unlabeled Drosera
“roundansticky” and Drosera capensis Complex. It also includes slightly
more difficult, subtropical species, such as Drosera filiformis ssp.
filiformis ‘Florida Red’ and Drosera capillaris, as well as difficult species,
such as Drosera regia. None of the species in this group goes truly
dormant and all are native to areas with warm conditions nearly year
round.
As with most other Drosera, most members of the tropical and
subtropical horticultural group have self-fertile flowers. Since many
of them grow year round, they can flower two or three times a year,
resulting in copious seed being produced. Inevitably, this seed is
blown into other pots throughout a grower’s collection, and quickly
become weeds, especially
members of the Drosera
Drosera madigascarensis
capensis Complex.
D. burmannii / D. sessilifolia
These two tiny, often confused, tropical Drosera are
remarkably similar looking. Drosera burmannii is native to Southeast
Asia and Australia. Drosera sessilifolia, in contrast, is native to South
America. Both Drosera burmannii and Drosera sessilifolia are tiny
plants with wedge shaped leaves, covered in fine sticky tentacles.
The end of the leaf has longer “snap” tentacles, which can move
rapidly towards the center of the plant, bringing prey in contact
with more tentacles. These “snap” tentacles can move within
seconds.
In cultivation, both of these annuals tend to become weeds, if
allowed to flower. Better color can be achieved by not feeding
them, although feeding is necessary for the plants to grow to full
size and flower. Powdered mealworms work well as food, but be
careful not to overwhelm these tiny plants with food particles that
Drosera burmannii
are too large.
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Drosera capensis ‘Pink’ in insufficient light.
D. capensis - Complex
The typical Drosera capensis is often the first sundew that a new carnivorous
plant enthusiast attempts to grow. It’s not hard to see why. This South African
native is a prolific seed producer, will grow in most any conditions, is virtually
indestructible, and a very showy species. It is also a highly variable species. At
present, the following “varieties” are known to exist: ‘Alba’, ‘Bains Kloof’, ‘Giant’,
‘Narrow Leaf’, ‘Pink’, ‘Red’, ‘Typical’, and ‘Wide Leaf’.
One of the most interesting is Drosera capensis ‘Pink’. This cultivar produces
leaves that are twice as large as regular Drosera capensis. The entire plant turns a light
pink under bright light, and has large, bright pink flowers. It is rumored that it is a
cross between Drosera capensis ‘Alba’ and Drosera capensis ‘Red’, but the true
parentage is unknown.
D. indica - Complex
One of the most widespread annual species is - or rather - was Drosera indica.
Recent work by Allen Lowrie and others has revealed that this, formerly variable
species, is actually a species complex, i.e. a series of closely related species. The Drosera hartmeyerorum
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CULTIVATING CARNIVOROUS PLANTS
Drosera indica Complex consists of the following species, with the following distributions:
Species
Range
Seeds
Notes
Near Kununurra, Western
Australia, and near Darwin,
Ovoid, 0.42 -
aquatica
Northwest Territory, Australia.
0.45 mm.
Affixed aquatic or free floating.
Prince Regent River to Drysdale
River in the Kimberley Region of
Ellipsoidal, 0.37
aurantiaca
Western Australia.
- 0.40 mm.
Plants red.
Prince Regent River to Drysdale
River in the Kimberley Region of
Ovoid, 0.31 -
Stem green with some red
barrettorum
Western Australia.
0.35 mm.
coloration, leaves green.
Prince Regent River to Wyndham
in the Kimberley Region of
Oblong, 0.86 -
cucullata
Western Australia.
0.90 mm.
Plant dull green, turning red.
Central Australia, China, Vietnam, Ellipsoidal, 0.50
Drier locations that seasonally
finlaysoniana
Laos, and Taiwan.
- 0.55 mm.
flood.
Emits a very strong honeydew
Broome to Darwin, Northwest
Broadly ovoid,
melon-like scent, possibly from
fragrans
Territory, Australia.
0.50 - 0.55 mm.
appendages near leaf bases.
Near Beverley Springs Station and Ovoid, 0.30 -
glabriscapa
Theda Station, Western Australia.
0.35 mm.
Plants green, turning red.
Broome to Keep River in the
Ovoid, 0.34 -
hartmeyerorum Northwest Territory of Australia.
0.35 mm.
Red stem and green-to-red leaves.
Africa, Madagascar, India, and
Ovoid, 0.4 - 0.5
indica
southeast Asia.
mm.
Plants red or green.
Ovoid, 0.4 - 0.5
Plants green with white flowers,
makinoi
Japan
mm.
often with pink anthers.
Short (3 in. (7 - 10 cm)), green
Near Darwin, Northwest
Ellipsoidal, 0.31
plants, having some red
nana
Territory, Australia.
- 0.33 mm.
coloration.
Northern Australia and southeast
Ovoid, 0.50 -
serpens
Asia.
0.55 mm.
Plants red or green.
Ovoid, 0.4 - 0.5
toyoakensis
Chubu District, Japan.
mm
Plants green with pink flowers.
As of now, only the members of the Drosera indica Complex from Australia and Japan have been studied
in-depth and placed into proper botanical categories. Members of the Drosera indica Complex from Africa,
Madigascar, India, and southeast Asia have yet to be studied in depth and are still labeled “Drosera indica.”
Most species of this complex in cultivation are Drosera finlaysoniana, although Drosera hartmeyerorum is
becoming more popular in cultivation. Most individuals in cultivation follow the typical annual-schedule of
requiring feeding when small plantlets, growing rapidly to become adults, and flowering profusely to set seed,
all within a few months.
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Drosera regia flower.
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D. regia
The King Sundew, Drosera regia, is one
of the rarest Drosera in the wild and most
desired Drosera in cultivation. Until recently,
it was rumored to be among the most
difficult to cultivate of the tropical Drosera.
Around the time of the registration of the
Drosera regia cultivar ‘Big Easy’ in 2003,
serious research began on the cultivation
and botanical history of the species. (‘Big
Easy’ does not require dormancy, does not
produce flowers, and is exceedingly, easy).
In the wild, Drosera regia has only every
been found at two locations, despite
extensive searches. Both sites are in the
Bainskloff Range near Wellington, South
Africa at elevations between 1,600 - 3,000 ft.
(500 - 900 m). The higher of the two sites
has since been overgrown by rushes and
other non-carnivorous species. The lower
site is in a similar state and has only a few
dozen plants remaining.
In its natural habitat, Drosera regia
grows in peaty quartzite sand with a gravel
cover along a seep formed along an old
creek bed. The habitat is maintained, like
many in the Fynbos, by fire periodically
sweeping through the area and burning off
larger, competing plants. The daytime
temperatures are mild, and nights are cool
(~60° F (15° C)). During the colder season,
when infrequent frosts occur, Drosera regia
Drosera regia
goes dormant, forming a tight cluster of
short, immature leaves.
Interestingly, Drosera regia shares many relic characteristics, such as woody rhizomes, not found in other
Drosera species. These characteristics set Drosera regia apart, and give context to early failures to cultivate the
species. Recent work - and study of the Drosera regia ‘Big Easy’ cultivar - has revealed that the species needs
food, a lot of food. Early growers often neglected to feed the plant after it germinated, resulting in its demise
due to starvation.
Drosera regia requires so much food, in fact, that it is advisable to include four or five Osmocote type,
slow release fertilizer pellets in the pure long-fiber sphagnum or mix of one part sphagnum peat to one part
sand and one part perlite. Drosera regia appears to prefer cooler roots, and less water than other Drosera. The
easiest method of propagation is via root cuttings. If given appropriate conditions, Drosera regia can produce
leaves up to 28 inches (70 cm) long.
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D. spathulata
The ubiquitous Drosera spathulata, native to
Southeast Asia and Australia, and contained in most
every carnivorous plant collection, is the default species
name for any round-and-sticky Drosera growing in
tropical or subtropical conditions. The plants may either
have white or pink flowers, will not produce
hibernaculum, and have been recorded to have
chromosome numbers 2n = 20, 2n = 30, and 2n = 40. It
would not be surprising to learn that a future, systematic
survey indicates that there are multiple species
comprising the Drosera spathulata complex.
So far, Drosera spathulata has been divided into the
following varieties - bakoensis (type location at Bako
National Park in Sarawak), gympiensis (type location near
Gympie, Australia), and lovellae (also known as “Fraser
Island Form”). The complex also includes Drosera
tokaiensis from Japan, which has a chromosome number
of 2n = 60, Drosera oblanceolata from China, and Drosera
neocaledonica from New Caledonia. Some varieties are
merely a half inch (2 cm) in diameter, while others, such
as var. lovellae, are 2 - 3 inches (up to 7 cm) in diameter.
Further, some are clearly tropical or subtropical variants,
Drosera spathulata
while others grow much better in alpine conditions.
D. tokaiensis
One of the most widespread and morphologically
variable species in cultivation is Drosera tokaiensis. Most likely,
this species is the result of the spread of fertile hybrid Drosera
x tokaiensis, which itself is a natural hybrid between Drosera
rotundifolia and Drosera spathulata. As discussed above, Drosera
spathulata is a, somewhat, morphologically variable species,
and occurs in numerous locations alongside Drosera
rotundifolia. Occasionally, the two hybridize. Given that
Drosera hybrids often share characteristics of both parents
and often look more like the mother plant, the hybrid
Drosera x tokaiensis is rather diverse in morphology. It is
unsurprising, then, that the species would be similarly
diverse. (The major difference between the hybrid and
Drosera natalensis, a species often
species is that the hybrid is 2n = 30 and the species is 2n =
confused with Drosera tokaiensis.
60).
As with most tropical and subtropical Drosera, this species is remarkably easy to grow, requiring only
warm temperatures, constant moisture, and light. It will do well outside in the summertime most anywhere in
the world. In the winter, in cold areas, it would be best to take it inside and grown on the tray method. Do
not be surprised, however, if it returns from seed after a cold winter outdoors.
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CULTIVATING CARNIVOROUS PLANTS
My standard growing guide for tropical and subtropical Drosera follows:
✵
Media: Very few tropical or subtropical Drosera demand a specific media. Most growers use a mix of
one part sphagnum peat to one part sand or perlite. Some growers have used pure sand for species
such as Drosera filiformis ssp. filiformis ‘Florida Red’, populations of which can be found naturally growing
in pure sand. Certain species, such as Drosera regia, should be potted in more specialized media, and
care should be taken to pot species from specialized environments in their appropriate media.
✵
Moisture: The easiest way to water most tropical and subtropical Drosera is via the tray method. Some
water should always be kept in the tray. Note, however, that in drier climates, this may not provide
sufficient humidity for the plants, some of which are used to very humid environments. Note also
that some species, such as Drosera regia, prefer drier growing conditions.
✵
Humidity: Most tropical and subtropical Drosera are not particularly picky about humidity, but some,
such as Drosera madigascarensis, may prefer more humid environments. Easy species, such as Drosera
capensis, can grow in almost any level of humidity and are a good gauge as to whether a particular
environment is humid enough for any carnivorous plants.
✵
Pot Size: Most species will do well in any pot larger than their diameter, but some, particularly those
with thick roots, should be given pots with ample room to grow. Generally, species with thicker
roots require larger pots.
✵
Feeding: As with most carnivorous plants, feeding is recommended. Note, however, that, especially
with tropical and subtropical Drosera, feeding will encourage flowering. Flowering will result in seed
production. Seed production will result in volunteer plantlets spreading outside the parent’s pot.
Generally speaking, the more a tropical or subtropical Drosera is fed, the more flowers it will produce.
Some, such as Drosera regia, do require supplemental feeding to survive, but most will capture so
many small insects on their own that fertilizer is not strictly necessary.
✵
Temperature: Tropical and subtropical Drosera should not be exposed to freezing temperatures for
long periods of time. Most will return from the roots if the aboveground portion is killed by frost,
but some are more temperamental. The upper limit for temperatures should not exceed 95° F (35°
C), if possible. If growing indoors, try to avoid temperatures that warm as it is unlikely that the plant
will be able to withstand such temperatures for long periods of time. Outside, most tropical and
subtropical Drosera can survive warmer temperatures in summer, but prolonged exposure can be
detrimental to long-term survival.
✵
Dormancy: Dormancy is not required for tropical or subtropical Drosera.
✵
Propagation: Tropical and subtropical Drosera are most easily propagated through seed, but can also
be propagated through leaf cuttings or root cuttings.
o
Seed: The easiest way to propagate tropical and subtropical Drosera is via seed. This is
produced in copious quantities for most species. The seed is mature when the pods dry, turn
black, and begin to split. Seed from neither tropical nor subtropical Drosera requires a cool
stratification period. It can be sown immediately and will germinate within weeks. Most
species will produce mature plants within months.
o
Leaf Cuttings: Leaf cuttings are a remarkably easy method of propagation. As with other
Drosera, cut the leaf from its base and place on an appropriate medium. In a few weeks, a
small plantlet should appear near one of the tentacles. It is beneficial to give the cutting even
higher humidity than the parent plants.
o
Root Cuttings: Perhaps the fastest way of propagating tropical and subtropical Drosera is via
root cuttings, but this only seems to work on thick-rooted species, such as Drosera capensis.
An inch long (2.5 centimeter) section of healthy root may be removed from a growing plant
and laid horizontally on top of a suitable media. The root should then be covered by a pinch
of media. If successful, growth should appear in a few weeks.
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Highland Tropical Drosera
Drosera latifolia
The highland tropical Drosera group includes a wide variety of species from the Tepuis of South
America, some cooler-growing species from South Africa, such as Drosera slackii, and the “Queensland
Sisters,” i.e. Drosera adelae, Drosera prolifera, and Drosera schizandra. All of these species are native to the highlands of their respective continents, and, although many of the characteristics of those highlands are very
different, the characteristic that unites all the members of this group is their need to grow in cool, humid
environments.
It may seem strange to think that the species of the lost worlds of South America have much in
common with the few species of the Fynbos which require cool temperatures or the inhabitants of Australia’s
rainforest, but, in cultivation, they are all remarkably similar. They would do well alongside Heliamphora or
highland Nepenthes.
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CULTIVATING CARNIVOROUS PLANTS
Drosera prolifera
D. adelae, D. prolifera, and D. schizandra - The Queensland Sisters
The so-called “Queensland Sisters”, the three species found
along stream banks in an area of coastal mountain rain forest in
northeast Queensland, Australia, are remarkably easy species to
grow, if given the right conditions. Drosera adelae is among the most
popular - and common - of the sisters. Although Drosera adelae will
tolerate virtually any condition, with the exception of hot sun
(above 90° F (32° C)) or freezing cold, it does best in semi-shade in
cool, humid areas.
Unlike Drosera adelae, Drosera prolifera and Drosera schizandra are
less tolerant of extreme conditions. Both prefer temperatures in-
line with those used to grow Heliamphora, and shade. Temperatures
should not exceed 75° F (24° C), humidity should be high, and
shade should be plentiful, especially for Drosera schizandra.
In recent years, hybrids among the Queensland Sisters have
come to the carnivorous plant market. The most desirable of these
is Drosera ‘Andromeda’, a hybrid between Drosera prolifera and
Drosera adelae ‘Red’
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NATCH GREYES
Drosera schizandra. Several different clones of this cross exist, and the more prolific ones have spread quite
widely in cultivation. As with many hybrids, Drosera ‘Andromeda’ exhibits a certain kind of vigor, making it as
easy to grow as Drosera adelae. As with the species, living sphagnum moss is the best media to grow Drosera
‘Andromeda’.
D. ascendens, esmeraldae, D. graomogolensis, latifolia, and other Tepui-dwellers
In the wild, most Tepui-dwelling
Drosera live among Heliamphora, and
advice on cultivation should be taken
from that chapter. As with Heliamphora
and other Tepui-dwellers, the Tepui-
dwelling Drosera, such as Drosera
ascendens, Drosera esmeraldae, and Drosera
graomogolensis, prefer cool temperatures
which never exceed 80° F (27° C), a
nighttime temperature drop of at least
10° F (5° C), high humidity, and bright
light during the day. Further, Tepui-
dwelling Drosera prefer deeper pots, as,
unlike their temperate cousins, Tepui-
dwellers have deep roots to anchor
themselves onto the rock and prevent
Drosera graomogolensis
their being swept away in the torrents of
water that follow a rainstorm.
In cultivation, many growers find success with a media mixture of one part sphagnum peat to one part
sand or pure, live sphagnum moss. Live sphagnum is preferable in growing environments where the ambient
humidity is lower than necessary for these species, as the sphagnum will generate some humidity itself during
its own photosynthetic cycle. Humidity should be as near as 100% as possible in order to mimic the humidity
naturally found atop the Tepuis. Daytime temperatures are best when they remain in the 65° - 70° F (18° -
21° C) range. Tepui-dwelling Drosera should be given as much light as possible.
D. slackii
This South African native grows in the cool, peaty wetlands
of the Klipriviersberg Mountains and the southern Kogelberg.
Although an easy plant in cultivation - and highly desirable - it is
not as common in cultivation as it once was, possibly as a factor
of many growers shifting focus towards Nepenthes, Heliamphora, and
other newly discovered carnivores.
In cultivation, Drosera slackii prefers a peat-based mix. Many
growers have reported success with a mix of equal parts
sphagnum peat and sand. Long-fiber sphagnum works well, too.
Cooler temperatures are necessary, as, when temperatures are too
warm, plants will exhibit obvious signs of heat stress such as
yellowing. High humidity environments are preferable. Drosera
Drosera slackii
slackii is easily propagated from leaf or root cuttings.
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CULTIVATING CARNIVOROUS PLANTS
My standard growing guide for highland tropical Drosera follows:
✵
Media: Most highland tropical Drosera are tolerant of a wide variety of mixtures, but most growers
use one part sphagnum peat and one part sand or long-fiber sphagnum. Some species, such as the
Queensland Sisters and some Tepui-dwelling Drosera, prefer living sphagnum. Tepui-dwelling Drosera
are, however, more temperamental about media and some experimentation may be necessary to find
the right blend.
✵
Moisture: As with most carnivores, highland tropical Drosera like moist, but not soaking media. A few
species, such as Drosera slackii, grow well using the traditional tray method, but others, such as Drosera
schizandra will not tolerate sitting in a tray of water for long periods of time.
✵
Humidity: Unlike some other horticultural categories of Drosera, all highland tropical Drosera come
from areas of high humidity. For many species in this category, insufficient humidity results in the
tentacles lacking “dew.” The solution to a lack of “dew” is to first assess the humidity level of the
environment. If increasing the humidity does not work, then a pest may be at work.
✵
Pot Size: Pots should be larger than the plant in diameter, but most are not particular about pot size
except the Tepui-dwelling Drosera. Those prefer deep pots.
✵
Feeding: Highland tropical Drosera, like most Drosera, may be fertilized or fed with small prey animals
(many of which can be purchased in the reptile section of pet stores). If using prey animals, be
careful to watch for leaf burn, which can occur much more easily on many of these species, if the
prey is too large or nutrient rich. Take particular care to watch Drosera schizandra for signs of leaf
burn, as that species has fewer sessile glands than most Drosera.
✵
Temperature: Highland tropical Drosera prefer the cooler temperatures experienced by Heliamphora or
highland Nepenthes. Daytime temperatures should not exceed 75° F (24° C). In the wild, most
experience night time temperatures from 50° - 60° F (10° - 16° C). Some species, such as Drosera
graomogolensis will tolerate warmer temperatures, but many species, such as Drosera ascendens require
cool temperatures for successful cultivation. In warmer temperatures, consider potting in live
sphagnum to increase the ambient humidity. This appears to give a cushion of about 5° F (2° C).
✵
Dormancy: Dormancy is not required for highland tropical Drosera.
✵
Propagation: Highland tropical Drosera can be propagated through seed, leaf cuttings, or root
cuttings.
o
Seed: The slowest way to propagate highland tropical Drosera is through seed. The seed does
not require a cool stratification period, but may be slow to germinate. For many species,
especially Tepui-dwellers, seed does not stay viable for long. Therefore, it is necessary to use
fresh seed, i.e. seed collected within a month or two of sewing. It will take a couple years for
a mature plant to result from seed sewn on a suitable medium.
o
Leaf Cuttings: Leaf cuttings are the easiest and most common method of propagation. As
with other Drosera, cut the leaf from its base and place on an appropriate medium. In a few
weeks, a small plantlet should appear near one of the tentacles. For Tepui-dwellers, leaf
cuttings have a lower success rate than with other Drosera. The Queensland Sisters and South
African Drosera, such as Drosera slackii, have, in contrast, a remarkably high success rate. It is
beneficial to give the cutting even higher humidity than the parent plant, and may be
beneficially to give them nearly as much light.
o
Root Cuttings: Perhaps the fastest way of propagating highland tropical Drosera is via root
cuttings, but this only seems to work on thick-rooted species or the Queensland Sisters.
Take an inch long (2.5 centimeter) section of healthy root from a healthy plant and lay it
horizontally on top of a suitable media. Cover this with a pinch of media, allowing it to be
exposed to light if using a root from the Queensland Sisters. In a few weeks, a plantlet
should sprout from somewhere along the root.
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NATCH GREYES
Drosera enodes setting seed after flowering.
Pygmy Drosera
The miniature jewels of Australia, the pygmy Drosera are tiny rosetted plants, usually less than an inch
(2.5 centimeter) in diameter. Most live in the southern portion of Western Australia. There, the climate is hot
and dry (~90° F (32° C)) in the summer and cool and wet (~46° F (8° C)) in the winter. As a consequence,
most species do most of their growing during the winter and go dormant during the summer.
The two species which live outside of Western Australia are Drosera pygmaea, which can be found
throughout Southern Australia as well as in New Zealand and Tasmania, and Drosera meristocaulis, which can
be found only in the a few valleys on the northern side of Pico da Neblina, the tallest mountain in Brazil.
Recent scientific analysis has revealed that Drosera meristocaulis (or, rather, its ancestor) was carried from
Australia by migrating birds sometime in the distant past.
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CULTIVATING CARNIVOROUS PLANTS
All species, except Drosera meristocaulis, produce gemmae.
These small plantlet buds are, by far, the easiest way to
reproduce pygmy Drosera. Once gathered from the crown of
the plant in the fall, they can be placed atop suitable media
and kept moist. In a few weeks, the gemmae will sprout into
new plants, genetic clones of their parents. Unfortunately, it is
not possible to store gemmae for very long, as they can easily
desiccate and die if kept too dry, or mold and rot if kept too
moist.
D. pulchella
Drosera pulchella is among is
easiest of pygmy Drosera to cultivate.
It forms a small rosette of tiny,
overlapping green petioles and
round, orange-colored traps. Among
the pygmies, it has the widest range
of flower colors, ranging from pink,
to white, to orange. As with many
pygmies, its gemmae are small, flat,
Drosera pulchella with gemmae.
and difficult to dislodge.
Drosera pulchella with
Drosera pulchella is tolerant of a wide-range of potting media, but prefers a sand-
a white flower.
based media, as do other pygmy Drosera. It will also tolerate a wide-range of
temperatures, growing from temperatures just above freezing to near 90° F (32° C).
D. scorpioides
The largest of the pygmy Drosera, Drosera
scorpioides, is among the easiest - and most popular - to
grow. As with all other pygmy Drosera, Drosera scorpioides
is a small, rosetted Drosera with small, thin petioles, and
large traps. Unlike other pygmies, new leaves grow
atop one another in a fashion that creates a stem. The
largest specimens of Drosera scorpioides can reach 4
inches (10 cm) tall, and be up to 2 inches (5 cm) in
diameter. The flowers are pink or white. In ideal
conditions, a single plant can live up to seven years,
and produce several hundred, if not thousand,
gemmae.
In the wild, Drosera scorpioides is found in deep,
white sand habitat in the Jarrah Forest region. At
times, temperatures can soar to over 110° F (45° C)!
(The heat reflecting white sand does keep the
temperature of the soil slightly lower, but not by
much).
Drosera scorpioides after gemmae removed.
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My standard growing guide for pygmy Drosera follows:
✵
Media: Most pygmy Drosera grow in a mixture which largely consists of silica sand in the wild. My
recommendation for cultivating these tiny sundews is to follow a soil recipe similar to the
composition found in the wild - three parts silica sand to one part sphagnum peat. Other successful
recipes include one part sand to one part sphagnum peat.
✵
Moisture: Pygmy Drosera, like many Australian carnivores, are adapted to long periods of dry weather
followed by long periods of wet weather. As with many Australian carnivores, pygmy Drosera grow
during the cool and wet season. They become dormant during the hot and dry season. In cultivation,
keep the media moist, but not wet, when the pygmy Drosera are actively growing. After gemmae have
formed, allow the media to dry out, but not completely. After a few months, increase the amount of
water available to the pygmy Drosera. This should induce growth. Keep cooler during active growth,
and hot during dormancy.
✵
Humidity: Humidity is not a major factor for pygmy Drosera, but ambient humidity must be above
30% for dew to form properly. If humidity is too low, pygmy Drosera will suffer.
✵
Pot Size: Most pygmy Drosera have deep roots and should be potted in deep pots. I have, however,
been successful at growing them in short, one-and-a-half inch (4 cm) pots, but I would not
recommend this set-up for long-term cultivation.
✵
Feeding: Pygmy Drosera must be fed in order to produce a healthy crop of gemmae and survive
through the hot, dormant period. Small, pulverized insects or diluted foliar fertilizer work well, but,
unlike some other carnivores, pygmy Drosera respond notably better to feeding with actual insects
than foliar fertilizer.
✵
Temperature: Pygmy Drosera are not particularly picky about temperatures, but will enter dormancy if
temperatures rise to 90° F (32° C). Active growth occurs best in temperatures around 70° F (21° C).
✵
Dormancy: Depending on the grower’s conditions, dormancy may or may not be required. High
temperatures will induce dormancy, which must be tolerated for several months. Keeping
temperatures low will allow the plants to grow year round without interruption. The problem with
keeping temperatures low, however, is that the plants will not form gemmae, which form in the
weeks before dormancy.
✵
Propagation: Pygmy Drosera are most easily propagated through gemmae, although it is possible to
propagate them through seed or leaf cuttings.
o
Gemmae: Gemmae are the easiest and best way to propagate pygmy Drosera. Gemmae are
formed after several months of active growth, just prior to dry dormancy. These asexual
“plantlets” form on the crown of the plant, clustering about the center. Most species
gemmae consists of semi-flat disks. These can be carefully detached from the crown of the
plant via gentle teasing with a pair of tweezers, spraying with water, or by using suction
created by gentle sucking on a straw (or creating a contraption with two straws and a
container which allows the gemmae to fall in the container). After harvesting, gemmae
should either be planted immediately or within the next couple weeks. They should be stored
in damp paper towels in a cool place. New plants should appear within a couple weeks of
sewing.
o
Seed: The slowest way to propagate pygmy Drosera is through seed. It is difficult to get seed
to germinate and, usually, requires up to a year of patience and long, alternating, cool damp
and hot dry periods.
o
Leaf Cuttings: Leaf cuttings are an uncommon way to propagate pygmy Drosera because the
leaves are so small. It is, however, possible, if a leaf can be detached with the petiole intact. It
should be placed atop a suitable media with a little media holding it down. In a few weeks, a
new plantlet should appear.
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CULTIVATING CARNIVOROUS PLANTS
Drosera peltata
Tuberous Drosera
The tuberous Drosera are an exclusively Australian phenomenon. Most
are native to Western Australia, a largely unexplored area of the country.
All die down to a tuber in the dry season, coming back when the rains
begin to wet the hard, dry earth. When growing, most form either rosettes
of large, broad leaves or tall, erect stems with many, roundish leaves.
The number of tuberous species is currently uncertain. In his recent
magnum opus, Allen Lowrie reported that he was aware of several more
species which have not yet been described. Undoubtedly, more have yet to
be found.
D. erythrorhiza
Drosera erythrorhiza is one of the largest, rosetted tuberous Drosera. Its
large, broad, green leaves are rimmed by carnivorous tentacles and can
grow up to 4 inches (10 cm) across. The center strip of leaf is largely
lacking in tentacles.
Drosera erythrorhiza13
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NATCH GREYES
As with other tuberous Drosera, Drosera erythrorhiza is best grown in well-drained, sandy soil in, at least, an
eight inch (20 cm) diameter pot. Secondary, lateral rhizomes are common with this species, and a good year’s
growth can result in a plethora of new plants coming up from all around the parent plant.
D. peltata - Complex
The Drosera peltata Complex consists of several
species of tuberous Drosera with long, scrambling stems
and shield-shaped leaves. Until recently, the entire
complex was known as the single, highly variable species
Drosera peltata. Recent taxonomic work has divided the
group into a number of species, including Drosera lunata
and Drosera peltata. At present, there is significant debate
about whether the widespread-species which covers
Southeast Asia is Drosera lunata or Drosera peltata.
Regardless of whether the plant is Drosera lunata or
Drosera peltata, the members of this complex are among
the easiest tuberous Drosera to cultivate. All will grow
rapidly, if fed small pulverized insects frequently. As with
all tuberous Drosera, feeding is necessary to allow large
tubers to grow. Like other tuberous Drosera, after a few
years of growth, most members of the Drosera peltata
complex will flower prolifically. These flowers are easily
pollinated and set copious amounts of seed, which will
easily germinate easily during the next “wet” season.
Drosera peltata
My standard growing guide for tuberous Drosera follows:
✵
Media: As with pygmy Drosera, most tuberous Drosera grow in a mixture which largely consists of
silica sand in the wild. My recommendation for cultivating these tiny sundews is to follow the natural
recipe of 3 parts silica sand to 1 part sphagnum peat.
✵
Moisture: Tuberous Drosera have a wet, active growth cycle, during which the media should be kept
wet, but not soaking, and a dry, dormancy cycle, during which the media should be kept near bone
dry.
✵
Humidity: Humidity is not a major factor, but keeping ambient humidity above 30% is important for
proper dew formation.
✵
Pot Size: Pots should be at least 6 inches (16 cm) deep, although deeper is better. Tubers can and
often will form outside the bottom of the pot if the pot is not large enough. Many species will form
horizontal tubers in good conditions, so relatively wide pots are recommended as well.
✵
Feeding: Tuberous Drosera must be fed in order to form tubers. Small, pulverized insects or diluted
foliar fertilizer are recommended.
✵
Temperature: Tuberous Drosera will enter dormancy if temperatures rise to 90° F (32° C). Active
growth occurs best in temperatures between 45° F (7° C) and 70° F (21° C).
✵
Dormancy: High temperatures will induce dormancy, and dormancy must occur for plants to survive
long-term in cultivation. During dormancy, the plants must be kept near bone dry for several months
in a warm location. Cool temperatures can induce growth, while too much water during dormancy
can root the tubers.
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CULTIVATING CARNIVOROUS PLANTS
✵
Propagation: Tuberous Drosera are most easily propagated through division of tubers, but can also be
propagated via seed.
o
Tuber Division: Many species will produce more than one tuber during the growing season,
if given plenty of food. Some, such as Drosera menziesii will produce more than one along the
main growing stem. Others, such as Drosera aberrans, will produce tubers along lateral
growths. During dormancy, these can be divided from one another and repotted, “eye” side
up.
o
Seed: The slowest way to propagate tuberous Drosera is through seed, although it is a fairly
easy process. Many species require heat stratification. For those species, if it is not possible
to leave them in an area which reaches 122° F (50° C), one can put the seeds in a tea bag and
submerge them in a pot of water at or near that temperature for a minute before sewing.
Alternatively, leave them sewn for a season or two. When conditions are right, they should
germinate.
o
Leaf Cuttings: Leaf cuttings have been reported as successful, but extremely rarely.
Petiolaris-Complex Drosera
Drosera ordensis14
Sometimes known as the “Woolly Sundews”, the leaf petioles of most of the Petiolaris-Complex are
covered in white hairs which protect them from the sun in the hot, dry locations where they live in their
native Australia. The hairs keep the leaves slightly cooler and lower the transpiration rate, as well as capture
some of the humidity in the air.
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Nearly all of the Petiolaris-Complex Drosera have small, round carnivorous leaves at the end of long, thin,
non-carnivorous petioles, reminiscent of Dionaea petioles. This makes them easily identifiable, and interesting
plants to grow.
The Petiolaris-Complex Drosera are native to an area of Australia that experiences warm temperatures year
round. The most difficult part about cultivating them is keeping them warm enough. They should experience
temperatures at or above 85° F (30° C), a will die if subjected to temperatures 65° F (18° C) or below.
Drosera falconeri15
D. falconeri
Drosera falconeri is one of the most desirable members of the Petiolaris-Complex. It grows a flat rosette
with a long petiole and large, flat leaf reminiscent of a Venus Flytrap’s trap. When it catches prey, its leaves
fold in half, much like a Venus Flytrap. Unlike other members of the Petiolaris-Complex, it grows in wetter
soil. The soil composition of its native habitat is sandy and alkaline. In cultivation, most growers are
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CULTIVATING CARNIVOROUS PLANTS
successful at using a mix of sphagnum peat and sand.
This species is among the most difficult of the complex to
cultivate successfully. Depending on the grower's conditions, it
can either be grown year round or subjected to a dry dormancy
season. If given a dry dormancy, the leaves will die back to a
tight rosette of leaves reminiscent of a hibernaculum. These
leaves will be gathered just under the soil surface.
D. petiolaris
Drosera petiolaris, the archetypical members of the Petiolaris-
Complex, is a rosetted plant that produces small, round traps
atop long, thin petioles. (Unlike some of its cousins, such as
Drosera derbyensis, Drosera petiolaris does not possess hairy
petioles). These traps are held out in a hemispherical pattern. In
strong light, the plants appear almost as if they were miniature
models of a solar-system.
The flowers of Drosera petiolaris, like the rest of the
Petiolaris-Complex, are self-incompatible. Two plants must be
flowering at the same time and cross-pollination occur for seed
to be produced. As with other species in the Petiolaris-Complex,
Drosera petiolaris readily hybridizes with other species within the
complex. Many plants in cultivation are, in fact, hybrids,
although not all are labeled as such. Therefore, it is necessary to
compare plants grown from seed against descriptions of the
species in order to ascertain whether the plants are, in fact, pure
species or hybrids.
Drosera lanata16
My standard growing guide for Petiolaris-Complex Drosera follows:
✵
Media: Petiolaris-Complex Drosera are not picky about potting media. A mixture of two parts sand to
one part sphagnum peat works well.
✵
Moisture: Petiolaris-Complex Drosera, like many Australian plants, have a have a wet, active growth
cycle and a drier, dormancy cycle. Unlike other Australian plants, the during the drier cycle, the
Petiolaris-Complex species do not need to have completely dry media. Instead, they may be kept just
barely moist.
✵
Humidity: Humidity is not a major factor, but keeping ambient humidity above 30% is important for
proper dew formation.
✵
Pot Size: Pots should be small, barely larger than the diameter of the plant, although slightly large
pots may be wanted if the plant is to slowly form clumps without requiring repotting.
✵
Feeding: Petiolaris-Complex Drosera should be fed with either small, pulverized insects or diluted foliar
fertilizer.
✵
Temperature: Unlike other carnivorous plant species, Petiolaris-Complex Drosera require warm
temperatures year round, as discussed above.
✵
Dormancy: Although Petiolaris-Complex Drosera do experience a dormancy, not much change is
necessary in terms of cultivation, merely reducing the amount of water available to the plant.
74
NATCH GREYES
✵
Propagation: Petiolaris-Complex Drosera will slowly form clumps over time, although seed is an easier
method of getting more plants faster. Leaf pullings can be done, but are difficult and not
recommended.
o
Division: For the species that produce clumps over time, it is recommended that dividing
these occurs during the drier dormancy season.
o
Seed: Easiest way to propagate Petiolaris-Complex Drosera is via seed, although it takes several
years to produce mature plants. Seed germinates readily without any special treatment, if
fresh. Temperatures should be kept warm for seedlings.
The following table of all known Drosera species includes their distribution and cultivation type in order
to facilitate thoughts on care:
Species
Distribution
Cultivation Type
aberrans
Australia
Tuberous
acaulis
South Africa
Tropical
adelae
Australia
Highland Tropical
admirabilis
South Africa
Subtropical
affinis
Tropical Africa
Tropical
afra
South Africa
Treat as Tuberous
alba
South Africa
Treat as Tuberous
aliciae
South Africa
Tropical
allantostigma
Australia
Pygmy
amazonica
Brazil
Highland Tropical
andersoniana
Australia
Tuberous
androsacea
Australia
Pygmy
anglica
North America, Europe, and Asia
Temperate
aquatica
Australia
Tropical
arcturi
Australia and New Zealand
Temperate
arenicola
Venezuela
Highland Tropical
ascendens
Brazil
Highland Tropical
aurantiaca
Australia
Tropical
auriculata
Australia
Tuberous
australis
Australia
Pygmy
banksii
Australia and New Guinea
Tuberous
barbigera ssp.
barbigera
Australia
Pygmy
barbigera ssp.
silvicola
Australia
Pygmy
barrettorum
Australia
Tropical
basifolia
Australia
Tuberous
bequaertii
Angola and Democratic Republic of the Congo
Tropical
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CULTIVATING CARNIVOROUS PLANTS
bicolor
Australia
Tuberous
binata
Australia and New Zealand
Subtropical/Temperate
bindoon
Australia
Pygmy
brevicornis
Australia
Petiolaris-Complex
brevifolia
North America, Central America, and South America
Subtropical
broomensis
Australia
Petiolaris-Complex
browniana
Australia
Tuberous
bulbigena
Australia
Tuberous
bulbosa
Australia
Tuberous
burkeana
Tropical Africa
Tropical
burmannii
Australia and Southeast Asia
Tropical
caduca
Australia
Petiolaris-Complex
callistos
Australia
Pygmy
calycina
Australia
Tuberous
camporupestris
Brazil
Highland Tropical
capensis
South Africa
Tropical
capillaris
North America, Central America, and South America
Subtropical
cayennensis
Brazil, French Guiana, and Venezuela
Subtropical
cendeensis
Venezuela
Highland Tropical
chrysolepis
Brazil and Peru
Highland Tropical
cistiflora
South Africa
Treat as Tuberous
citrina
Australia
Pygmy
closterostigma
Australia
Pygmy
coalara
Australia
Pygmy
coccipetala
South Africa
Treat as Tuberous
collina
Australia
Tuberous
collinsiae
South Africa
Tropical
communis
Brazil, Colombia, Paraguay, and Venezuela
Highland Tropical
coomallo
Australia
Pygmy
cucullata
Australia
Tropical
cuneifolia
South Africa
Treat as Tuberous
darwinensis
Australia
Petiolaris-Complex
depauperata
Australia
Pygmy
derbyensis
Australia
Petiolaris-Complex
dichrosepala
Australia
Pygmy
dielsiana
Southern Africa
Tropical
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NATCH GREYES
dilatatopetiolaris
Australia
Petiolaris-Complex
drummondii
Australia
Tuberous
echinoblastus
Australia
Pygmy
elongata
Angola
Tropical
eneabba
Australia
Pygmy
enodes
Australia
Pygmy
eremaea
Australia
Tuberous
ericgreenii
South Africa
Treat as Tuberous
erythrogyne
Australia
Tuberous
erythrorhiza
Australia
Tuberous
esmeraldae
Colombia, Venezuela
Highland Tropical
esperensis
Australia
Tuberous
esterhuyseniae
South Africa
Tropical
falconeri
Australia
Petiolaris-Complex
felix
Venezuela
Highland Tropical
filiformis ssp.
filiformis
North America
Temperate
filiformis ssp. tracyi
Florida
Subtropical
fimbriata
Australia
Tuberous
finlaysonia
Australia
Tropical
fragrans
Australia
Tropical
fulva
Australia
Petiolaris-Complex
geniculata
Australia
Tuberous
gibsonii
Australia
Pygmy
gigantea
Australia
Tuberous
glabripes
South Africa
Subtropical
glabriscapa
Australia
Tropical
Treat as Highland
glanduligera
Australia
Tropical
gracilis
Australia
Tuberous
graminifolia
Brazil
Highland Tropical
graniticola
Australia
Tuberous
grantsaui
Brazil
Highland Tropical
graomogolensis
Brazil
Highland Tropical
grievei
Australia
Pygmy
hamiltonii
Australia
Subtropical
hartmeyerorum
Australia
Tropical
helodes
Australia
Pygmy
77
CULTIVATING CARNIVOROUS PLANTS
heterophylla
Australia
Tuberous
hilaris
South Africa
Treat as Tuberous
hirsuta
Australia
Tuberous
hirtella
Brazil
Highland Tropical
hirticalyx
Venezuela
Highland Tropical
hookeri
Australia
Tuberous
huegelii
Australia
Tuberous
humbertii
Madagascar
Tropical
humilis
Australia
Tuberous
hyperostigma
Australia
Pygmy
indica
Tropical Africa and Southeast Asia
Tropical
indumenta
Australia
Tuberous
Temperate Europe, Temperate North America,
intermedia
Tropical South America
Temperate/Tropical
intricata
Australia
Tuberous
kaieteurensis
Guyana, Trinidad and Tobago, and Venezuela
Highland Tropical
katangensis
Democratic Republic of the Congo
Tropical
kenneallyi
Australia
Petiolaris-Complex
lanata
Australia
Petiolaris-Complex
lasiantha
Australia
Pygmy
leioblastus
Australia
Pygmy
leucoblasta
Australia
Pygmy
leucostigma
Australia
Pygmy
linearis
North America
Temperate
longiscapa
South Africa
Tropical
lowriei
Australia
Tuberous
lunata
Australia
Tuberous
macrantha
Australia
Tuberous
macrophylla
Australia
Tuberous
madagascariensis
Africa
Tropical
magna
Australia
Tuberous
major
Australia
Tuberous
makinoi
Japan
Tropical
mannii
Australia
Pygmy
marchantii
Australia
Tuberous
menziesii
Australia
Tuberous
meristocaulis
Venezuela
Pygmy
78
NATCH GREYES
micra
Australia
Pygmy
microantha
Australia
Pygmy
microphylla
Australia
Tuberous
microscapa
Australia
Pygmy
miniata
Australia
Pygmy
minutiflora
Australia
Pygmy
modesta
Australia
Tuberous
monantha
Australia
Tuberous
montana
Argentina, Brazil, Paraguay, and Venezuela
Highland Tropical
monticola
Australia
Tuberous
moorei
Australia
Tuberous
murfetii
Australia
Temperate
myriantha
Australia
Tuberous
nana
Australia
Tropical
natalensis
Madagascar, Mozambique, South Africa
Tropical
neesii
Australia
Tuberous
neocaledonica
New Caledonia
Tropical
nidiformis
South Africa
Tropical
nitidula
Australia
Pygmy
nivea
Australia
Pygmy
oblanceolata
Hong Kong
Subtropical
occidentalis
Australia
Pygmy
omissa
Australia
Pygmy
orbiculata
Australia
Tuberous
ordensis
Australia
Petiolaris-Complex
oreopodion
Australia
Pygmy
paleacea
Australia
Pygmy
pallida
Australia
Tuberous
paradoxa
Australia
Petiolaris-Complex
patens
Australia
Pygmy
pauciflora
South Africa
Treat as Tuberous
pedicellaris
Australia
Pygmy
peltata
Southeast Asia
Tuberous
peruensis
Peru
Highland Tropical
petiolaris
Australia, New Guinea
Petiolaris-Complex
pilosa
Cameroon, Guinea, Kenya, Tanzania
Tropical
planchonii
Australia
Tuberous
79
CULTIVATING CARNIVOROUS PLANTS
platypoda
Australia
Tuberous
platystigma
Australia
Pygmy
porrecta
Australia
Tuberous
praefolia
Australia
Tuberous
prolifera
Australia
Highland Tropical
prophylla
Australia
Tuberous
prostrata
Australia
Tuberous
prostratoscaposa
Australia
Tuberous
pulchella
Australia
Pygmy
purpurascens
Australia
Tuberous
pycnoblasta
Australia
Pygmy
pygmaea
Australia, New Zealand
Pygmy
quartzicola
Brazil
Highland Tropical
radicans
Australia
Tuberous
ramellosa
Australia
Tuberous
ramentacea
South Africa
Treat as Tuberous
rechingeri
Australia
Pygmy
regia
South Africa
Subtropical
roraimae
Brazil, Guyana, Venezuela
Highland Tropical
roseana
Australia
Pygmy
rosulata
Australia
Tuberous
rotundifolia
North America, Europe, Asia
Temperate
rubrifolia
South Africa
Subtropical
rupicola
Australia
Tuberous
salina
Australia
Tuberous
sargentii
Australia
Pygmy
schizandra
Australia
Highland Tropical
schmutzii
Australia
Tuberous
schwackei
Brazil
Highland Tropical
scorpioides
Australia
Pygmy
serpens
Australia
Tropical
sessilifolia
Brazil, Guyana, Venezuela
Tropical
sewelliae
Australia
Pygmy
silvicola
Australia
Pygmy
slackii
South Africa
Subtropical
solaris
Guyana
Highland Tropical
spatulata
Australia, New Zealand, Southeast Asia
Subtropical
80
NATCH GREYES
spilos
Australia
Pygmy
spiralis
Brazil
Highland Tropical
squamosa
Australia
Tuberous
stelliflora
Australia
Pygmy
stenopetala
New Zealand
Temperate
stolonifera
Australia
Tuberous
stricticaulis
Australia
Tuberous
subhirtella
Australia
Tuberous
subtilis
Australia
Tuberous
sulphurea
Australia
Tuberous
thysanosepala
Australia
Tuberous
tentaculata
Brazil
Highland Tropical
tokaiensis
Japan
Subtropical
tomentosa
Brazil
Highland Tropical
toyoakensis
Japan
Tropical
trichocaulis
Australia
Pygmy
trinervia
South Africa
Treat as Tuberous
tubaestylis
Australia
Tuberous
ultramafica
Southeast Asia
Highland Tropical
uniflora
Argentina, Chile, and Falkland Islands
Temperate
venusta
South Africa
Tropical
verrucata
Australia
Pygmy
villosa
Brazil
Highland Tropical
viridis
Brazil
Highland Tropical
walyunga
Australia
Pygmy
whittakeri
Australia
Tuberous
yilgarnensis
Australia
Tuberous
yutajensis
Venezuela
Highland Tropical
zeyheri
South Africa
Treat as Tuberous
zigzagia
Australia
Tuberous
zonaria
Australia
Tuberous
81